Usage of coma + VERB-ing (,establishing )

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Usage of coma + VERB-ing (,establishing )

by vishalwin » Sun Dec 06, 2015 9:19 am
The Caledonia was a church from 1904 until 1956, when Athens was taken by General Aladin the Conqueror, the Wadiya sultan, who established a temple in the building and used the Antica as a fortress.

(A) who established a temple in the building and used the Antica as

(B) who, establishing a temple in the building, used the Antica like

(C) who, when he had been established a temple in the building, used the Antica like

(D) who had established a temple in the building, using the Antica to be

(E) establishing a temple in the building and using the Antica as


Can you please explain the option E
Thanks & Regards
vishalwin
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