Challenging the hypothesis that tuberculosis could only be transmitted when someone inhaled air that had been exhaled by an infected individual, an article published in Nature magazine in 1872 reported that when examined, monkeys that had been kept in confinement and had consequently never breathed contaminated air still displayed tubercles in their lungs and presented with the same symptoms that individuals infected with tuberculosis did, and that they did not always transmit the disease upon coming in contact with an uninfected monkey.

(A) and had consequently never breathed contaminated air still displayed tubercles in their lungs and presented with the same symptoms that individuals infected with tuberculosis did, and that

(B) consequently never breathed contaminated air, still displaying tubercles in their lungs and presenting with the same symptoms as individuals who had been infected with tuberculosis did, and that

(C) and consequently had never breathed contaminated air, still displayed tubercles in their lungs and had presented the same symptoms as individuals infected with tuberculosis, and

(D) had consequently never breathed contaminated air, but still they displayed tubercles in their lungs and presented with the same symptoms as individuals infected with tuberculosis, and that

(E) and consequently they had never breathed contaminated air still displayed tubercles in their lungs and presented with the same symptoms that individuals infected with tuberculosis, and