Usage of coma + VERB-ing (,establishing )

[vishalwin]

The Caledonia was a church from 1904 until 1956, when Athens was taken by General Aladin the Conqueror, the Wadiya sultan, who established a temple in the building and used the Antica as a fortress.

(A) who established a temple in the building and used the Antica as

(B) who, establishing a temple in the building, used the Antica like

(C) who, when he had been established a temple in the building, used the Antica like

(D) who had established a temple in the building, using the Antica to be

(E) establishing a temple in the building and using the Antica as


Can you please explain the option E