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List matches with the verb has - both singular.
Animals matches with the verb exhibit - both plural.
Here's the structure of the sentence:
The list - of animals - that exhibit X - has been expanded to include Y
Main subj - prep phrase - "that" modifier, modifying the preceding noun, animals - main verb (plus the rest of the sentence)
Read it as:
The list (of animals that exhibit X) has been expanded (to include Y).
You can also use logic to help you out a little. Are the animals exhibiting X or is the list exhibiting X? X is a characteristic of animals, not of a list - a list can't exhibit a preference for left or right-handedness.
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Stacey Koprince
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