Studies of the human "sleep-wake cycle" have practical relevance for matters ranging from duty assignments in nuclear submarines and air-traffic control towers to the staff of shifts in 24-hour factories.
(A) to the staff of
(B) to those who staff
(C) to the staffing of
(D) and staffing
(E) and the staff of
(A) to the staff of
(B) to those who staff
(C) to the staffing of
(D) and staffing
(E) and the staff of
