Sentence Correction

In a crowded, acquisitive world, the disappearance of lifestyles such as those once followed by southern Africa's Bushmen and Australia's aborigines, requiring vast wild spaces and permitting little accumulation of goods, seem inevitably doomed.

(A) requiring vast wild spaces and permitting little accumulation of goods, seem inevitably doomed

(B) requiring vast wild spaces and permitting little accumulation of goods, seems to be inevitably doomed

(C) which require vast wild spaces and permit little accumulation of goods, seems to be inevitably doomed

(D) life-styles that require vast wild spaces and permit little accumulation of goods, seem inevitable

(E) life-styles requiring vast wild spaces and permitting little accumulation of goods, seems inevitable

[spoiler]
This question is in Sahil's grammar notes available in BTG forum.OA later after discussions.
One doubt here seem/seems modifies what?[/spoiler]