Although when a hagfish is threatened, it will secrete slime that is small in quantity, it expands several hundred times as it absorbs seawater, forming a slime ball that can coat the gills of predatory fish and either suffocate them or distress them enough to make them flee.

A. Although when a hagfish is threatened, it will secrete slime that is small in quantity,
B. Although a small quantity of slime is secreted by the hagfish, when threatened
C. Although, when threatened, a hagfish will secrete slime that is small in quantity,
D. Although the slime secreted by a threatened hagfish is small in quantity,
E. Although the hagfish secretes a small quantity of slime when threatened,

D