A new genetically engineered papaya was produced not by seed companies
who have been motivated by profit, which was the case with most genetically
modified crops previously approved for commercial use, but university
and United States Department of Agriculture researchers who allowed growers
to use it free of charge.

(A) produced not by seed companies who have been motivated by profit, which
was the case with most genetically modified crops previously approved for commercial use, but

(B) produced not by profit-motivated seed companies, the case with most genetically
modified crops previously approved for commercial use, but

(C) produced not by profit-motivated seed companies, as was the case with most
genetically modified crops previously approved for commercial use, but by

(D) not produced by profit-motivated seed companies, which was the case with
most genetically modified crops previously approved for commercial use, but was produced by

(E) not produced by seed companies who have been motivated by profit, the case with
most genetically modified crops previously approved for commercial use, but by

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