In 1929 relatively small declines in the market ruined many speculators having bought Wed margin; they had to sell, and their selling pushed other investors to the brink.

(A) specu1ators having bought Wed margin; they had to sell, and
(B) speculators who had bought Wed margin, having had to sell,
(C) speculators who had bought Wed margin; they had to sell, and
(D) speculators, those who bad bought Wed margin; these speculators had to sell, and
(E) speculators, who, having bought Wed margin and having to sell,