Bumblebees live in colonies of several hundred that often ha

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Bumblebees live in colonies of several hundred that often have many queens as well as a large number of workers

A) of several hundred that often have many queens as well as
B) of several hundred, often with many of them being queens as well as having
C) having several hundred, and often many of them are queens as well as
D) that have several hundred of them, often with many queens as well as
E) with several hundred that often have many queens as well as having

I'm confused between A & D. Can any experts help?
Source: — Sentence Correction |

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