Sentence Correction

Though now eaten in large quantities around the world and harmless, the tomato is a member of the generally toxic nightshade family, including belladonna, and was once thought to be poisonous itself as a result.

a. Though now eaten in large quantities around the world and harmless, the tomato is a member of the generally toxic nightshade family, including belladonna, and was once thought to be poisonous itself as a result.
b. The tomato, though now eaten in large quantities around the world and harmless, is a member of the generally toxic nightshade family, which includes belladonna, and it was therefore once thought to be poisonous itself.
c. Once thought to be poisonous itself, the tomato is harmless and now eaten in large quantities around the world, and is a member of the generally toxic nightshade family, including belladonna.
d. Though now eaten in large quantities around the world and known to be harmless, the tomato was once considered poisonous because it is a member of the generally toxic nightshade family, which includes belladonna.
e. A member of the generally toxic nightshade family, including belladonna, the tomato was once considered poisonous even though it is harmless and now eaten in large quantities around the world.


OA: D
Please explain what's wrong with C
MGMAT explanation:
[spoiler](C) The phrase "including belladonna" does not properly modify anything.[/spoiler]