The Parthenon was a church from 1204 until 1456, when Athens

The Parthenon was a church from 1204 until 1456, when Athens was taken by General Mohammed the Conqueror, the Turkish sultan, who established a mosque in the building and used the Acropolis as a fortress.

(A) who established a mosque in the building and used the Acropolis as

(B) who, establishing a mosque in the building, used the Acropolis like

(C) who, when he had established a mosque in the building, used the Acropolis like

(D) who had established a mosque in the building, using the Acropolis to be

(E) establishing a mosque in the building and using the Acropolis as

OA A

Source: Official Guide

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