OG: Upon their first encountering leaf-cutting ants in South America, the insects

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Upon their first encountering leaf-cutting ants in South America, the insects seemed to some Europeans to be carrying bits of greenery to shade themselves from the tropical sun – hence the sobriquet “parasol ants.”

A. Upon their first encountering leaf-cutting ants in South America, the insects seemed to some Europeans to be
B. Upon their first encountering leaf-cutting ants in South America, the insects seemed to some Europeans to be
C. On first encountering leaf-cutting ants in South America, it seemed to some Europeans that the insects were
D. On first encountering leaf-cutting ants in South America, some Europeans thought the insects were
E. On their first encounter with leaf-cutting ants in South America, some Europeans thought it was because the insects were

D